Some acciaccature don't play back despite being marked to

Bar 424 and onward in the attached score, trumpet alto sax tenor sax trombone
I tried deleting and manually reinserting (since there had been copypaste & explode shenanigans) and it didn't change.
It isn't the usual "musescore starts dropping notes after a while" issue because shutting down the program and relaunching doesn't make them play again.
It isn't grace notes in general because it works in other points in the same score.